nydus/Contributions to the Theory of Natural SelectionPublic
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IV. THE MALAYAN PAPILIONIDÆ OR SWALLOW-TAILED BUTTERFLIES, AS ILLUSTRATIVE OF THE THEORY OF NATURAL SELECTION.
Co-existing Variety.—This is a somewhat doubtful case. It is when
a slight but permanent and hereditary modification of form exists in
company with the parent or typical form, without presenting those
intermediate gradations which would constitute it a case of simple
variability. It is evidently only by direct evidence of the two forms
breeding separately that this can be distinguished from dimorphism. The
difficulty occurs in Papilio Jason, and P. Evemon, which inhabit the
same localities, and are almost exactly alike in form, size, and
colouration, except that the latter always wants a very conspicuous red
spot on the under surface, which is found not only in P. Jason, but in
all the allied species. It is only by breeding the two insects that it
can be determined whether this is a case of a co-existing variety or of
dimorphism. In the former case, however, the difference being constant
and so very conspicuous and easily defined, I see not how we could
escape considering it as a distinct species. A true case of co-existing
forms would, I consider, be produced, if a slight variety had become
fixed as a local form, and afterwards been brought into contact with the
parent species, with little or no intermixture of the two; and such
instances do very probably occur.
Race or subspecies.—These are local forms completely fixed and isolated; and there is no possible test but individual opinion to determine which of them shall be considered as species and which varieties. If stability of form and “*the constant transmission of some characteristic