- The Origin of Species (pp. 1–2)
- An Historical Sketch of the Progress of Opinion on the Origin of Species, Previously to the Publication of the First Edition of This Work (pp. 3–20)
- II (pp. 21–24)
- Individual Differences (pp. 25–38)
- Species of the Larger Genera in Each Country Vary More Frequently Than the Species of the Smaller Genera (pp. 39–43)
- Summary (p. 44)
- III (pp. 45–47)
- Geometrical Ratio of Increase (pp. 48–55)
- Complex Relations of All Animals and Plants to Each Other in the Struggle for Existence (pp. 56–64)
- V (pp. 65–73)
- Acclimatisation (pp. 74–77)
- Correlated Variation (pp. 78–82)
- A Part Developed in Any Species in an Extraordinary Degree or Manner, in Comparison with the Same Part in Allied Species, Tends to Be Highly Variable (pp. 83–87)
- Specific Characters More Variable Than Generic Characters (pp. 88–91)
- Distinct Species Present Analogous Variations, So That a Variety of One Species Often Assumes a Character Proper to an Allied Species, or Reverts to Some of the Characters of an Early Progenitor (pp. 92–101)
- Summary (pp. 102–103)
- VI (p. 104)
- On the Absence or Rarity of Transitional Varieties (pp. 105–147)
- Summary: The Law of Unity of Type and of the Conditions of Existence Embraced by the Theory of Natural Selection (pp. 148–151)
- VII (pp. 152–206)
- VIII (pp. 207–212)
- Inherited Changes of Habit or Instinct in Domesticated Animals (pp. 213–216)
- Special Instincts (pp. 217–251)
- XIII (pp. 252–261)
- Absence of Batrachians and Terrestrial Mammals on Oceanic Islands (pp. 262–274)
- Summary of the Last and Present Chapters (pp. 275–278)
- XV (pp. 279–313)
- Glossary of the Principal Scientific Terms Used in the Present Volume (pp. 314–333)
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