Accordingly all natural arrangements, whether the reality of the distinction of Kinds was felt or not by their framers, have been led, by the mere pursuit of their own proper end, to conform themselves to the distinctions of Kind, so far as these have been ascertained at the time. The species of Plants are not only real Kinds, but are probably, all of them, real lowest Kinds, Infimæ Species; which, if we were to subdivide, as of course it is open to us to do, into sub-classes, the subdivision would necessarily be founded on definite distinctions, not pointing (apart from what may be known of their causes or effects) to any difference beyond themselves.
In so far as a natural classification is grounded on real Kinds, its groups are certainly not conventional: it is perfectly true that they do not depend upon an arbitrary choice of the naturalist. But it does not follow, nor, I conceive, is it true, that these classes are determined by a type, and not by characters. To determine them by a type would be as sure a way of missing the Kind, as if we were to select a set of characters arbitrarily. They are determined by characters, but these are not arbitrary. The problem is, to find a few definite characters which point to the multitude of indefinite ones. Kinds are Classes between which there is an impassable barrier; and what we have to seek is, marks whereby we may determine on which side of the barrier an object takes its place. The characters which will best do this should be chosen: if they are also important in themselves, so much the better. When we have selected the characters, we parcel out the objects according to those characters, and not, I conceive, according to resemblance to a type. We do not compose the species Ranunculus acris, of all plants which bear a satisfactory degree of resemblance to a model buttercup, but of those which possess certain characters selected as marks by which we might recognize the possibility of a common parentage; and the enumeration of those characters is the definition of the species.
The question next arises, whether, as all Kinds must have a place among the classes, so all the classes in a natural arrangement must be Kinds? And to this I answer, certainly not. The distinctions of Kinds are not numerous enough to make up the whole of a classification. Very few of the genera of plants, or even of the families, can be pronounced with certainty to be Kinds. The great distinctions of Vascular and Cellular, Dicotyledonous or Exogenous and Monocotyledonous or Endogenous plants, [pg 504] are perhaps differences of kind; the lines of demarcation which divide those classes seem (though even on this I would not pronounce positively) to go through the whole nature of the plants. But the different species of a genus, or genera of a family, usually have in common only a limited number of characters. A Rose does not seem to differ from a Rubus, or the Umbelliferæ from the Ranunculaceæ, in much else than the characters botanically assigned to those genera or those families. Unenumerated differences certainly do exist in some cases; there are families of plants which have peculiarities of chemical composition, or yield products having peculiar effects on the animal economy. The Cruciferæ and Fungi contain an unusual proportion of nitrogen; the Labiatæ are the chief sources of essential oils, the Solaneæ are very commonly narcotic, etc. In these and similar cases there are possibly distinctions of Kind; but it is by no means indispensable that there should be. Genera and Families may be eminently natural, though marked out from one another by properties limited in number; provided those properties are important, and the objects contained in each genus or family resemble each other more than they resemble any thing which is excluded from the genus or family.
After the recognition and definition, then, of the infimæ species, the next step is to arrange those infimæ species into larger groups: making these groups correspond to Kinds wherever it is possible, but in most cases without any such guidance. And in doing this it is true that we are naturally and properly guided, in most cases at least, by resemblance to a type. We form our groups round certain selected Kinds, each of which