often range widely, and as if capriciously; for in two adjoining river systems some of the species may be the same and some wholly different.
It is probable that they are occasionally transported by what may be called accidental means. Thus fishes still alive are not very rarely dropped at distant points by whirlwinds; and it is known that the ova retain their vitality for a considerable time after removal from the water. Their dispersal may, however, be mainly attributed to changes in the level of the land within the recent period, causing rivers to flow into each other. Instances, also, could be given of this having occurred during floods, without any change of level. The wide differences of the fish on the opposite sides of most mountain-ranges, which are continuous and consequently must, from an early period, have completely prevented the inosculation of the river systems on the two sides, leads to the same conclusion. Some freshwater fish belong to very ancient forms, and in such cases there will have been ample time for great geographical changes, and consequently time and means for much migration. Moreover, Dr. Gunther has recently been led by several considerations to infer that with fishes the same forms have a long endurance. Saltwater fish can with care be slowly accustomed to live in fresh water; and, according to Valenciennes, there is hardly a single group of which all the members are confined to fresh water, so that a marine species belonging to a freshwater group might travel far along the shores of the sea, and could, it is probable, become adapted without much difficulty to the fresh waters of a distant land.
Some species of freshwater shells have very wide ranges, and allied species which, on our theory, are descended from a common parent, and must have proceeded from a single source, prevail throughout the world. Their distribution at first perplexed me much, as their ova are not likely to be transported by birds; and the ova, as well as the adults, are immediately killed by seawater. I could not even understand how some naturalised species have spread rapidly throughout the same country. But two facts,