fenestra ovalis, or oval window. As liquids are practically incompressible, nature has made allowance for the squeezing in of the oval window membrane, by providing a second opening, the round window, also covered with a membrane. When the stapes pushes the oval membrane in, the round membrane bulges out, its elasticity sufficing to put a certain pressure on the perilymph (indicated by the dotted portion of the inner ear).
The inner ear consists of two main parts, the cochlea—so called from its resemblance in shape to a snail's shell—and the semicircular canals. Each portion has its perilymph and endolymph, and contains a number of the nerve-ends, which are, however, most numerous in the cochlea. We do not know for certain what the functions of the canals and the cochlea are; but it is probable that the former enables us to distinguish between the intensity or loudness of sounds and the direction from which they come, while the latter enables us to determine the pitch of a note. In the cochlea are about 2,800 tiny nerve-ends, called the rods of Corti. The normal ear has such a range as to give about 33 rods to the semitone. The great scientist Helmholtz has advanced the theory that these little rods are like tiny tuning-forks, each responding to a note of a certain pitch; so that when a string of a piano is sounded and the air vibrations are transmitted to the inner ear, they affect only one of these rods and the part of the brain which it serves, and we have the impression of one particular note. It has been proved by experiment that a very sensitive ear can distinguish between sounds varying in pitch by only 1⁄64th of a semitone, or but half the range