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nydus/Contributions to the Theory of Natural SelectionPublic

This volume collects essays written by Alfred Russel Wallace regarding his independent discovery and development of the theory of natural selection. The author provides corrected and expanded versions of his previous scientific writings to demonstrate the scope and application of his findings.

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IV. THE MALAYAN PAPILIONIDÆ OR SWALLOW-TAILED BUTTERFLIES, AS ILLUSTRATIVE OF THE THEORY OF NATURAL SELECTION.

of some characteristic peculiarity of organization,” constitutes a species. Now leaving out the question of “origin,” which we cannot determine, and taking only the proof of separate origin, “the constant transmission of some characteristic peculiarity of organization,” we have a definition which will compel us to neglect altogether the amount of difference between any two forms, and to consider only whether the differences that present themselves are permanent*. The rule, therefore, I have endeavoured to adopt is, that when the difference between two forms inhabiting separate areas seems quite constant, when it can be defined in words, and when it is not confined to a single peculiarity only, I have considered such forms to be species. When, however, the individuals of each locality vary among themselves, so as to cause the distinctions between the two forms to become inconsiderable and indefinite, or where the differences, though constant, are confined to one particular only, such as size, tint, or a single point of difference in marking or in outline, I class one of the forms as a variety of the other.

I find as a general rule that the constancy of species is

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