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nydus/Contributions to the Theory of Natural SelectionPublic

This volume collects essays written by Alfred Russel Wallace regarding his independent discovery and development of the theory of natural selection. The author provides corrected and expanded versions of his previous scientific writings to demonstrate the scope and application of his findings.

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Table of Contents

IV. THE MALAYAN PAPILIONIDÆ OR SWALLOW-TAILED BUTTERFLIES, AS ILLUSTRATIVE OF THE THEORY OF NATURAL SELECTION.

Papilio Agamemnon exhibits a somewhat similar series of changes. In India it is always tailed; in the greater part of the archipelago it has a very short tail; while far east, in New Guinea and the adjacent islands, the tail has almost entirely disappeared.

In the Polydorus-group two species, P. Antiphus and P. Diphilus, inhabiting India and the Indian region, are tailed, while the two which take their place in the Moluccas, New Guinea, and Australia, P. Polydorus and P. Leodamas, are destitute of tail, the species furthest east having lost this ornament the most completely.

Western species, Tailed.Allied Eastern species not Tailed.
Papilio Pammon (India)P. Thesus (Islands) minute tail.
P. Agamemnon, var. (India)P. Agamemnon, var. (Islands).
P. Antiphus (India, Java)P. Polydorus (Moluccas).
P. Diphilus (India, Java)P. Leodamas (New Guinea).

The most conspicuous instance of local modification of form, however, is exhibited in the island of Celebes, which in this respect, as in some others, stands alone and isolated in the whole archipelago. Almost every species of Papilio inhabiting Celebes has the wings of a peculiar shape, which distinguishes them at a glance from the allied species of every other island. This peculiarity consists, first, in the upper wings being generally more elongate and falcate; and secondly, in the costa or anterior margin being much more curved, and in most instances exhibiting near the base an abrupt bend or elbow, which in some species is very conspicuous. This peculiarity is visible, not only when the Celebesian species are compared with their small-sized allies of Java and Borneo, but also, and in an almost equal degree, when the large forms of Amboyna and the Moluccas are the objects of comparison, showing that this is quite a distinct phenomenon from the difference of size which has just been pointed out.

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