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nydus/Contributions to the Theory of Natural SelectionPublic

This volume collects essays written by Alfred Russel Wallace regarding his independent discovery and development of the theory of natural selection. The author provides corrected and expanded versions of his previous scientific writings to demonstrate the scope and application of his findings.

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Table of Contents

I. ON THE LAW WHICH HAS REGULATED THE INTRODUCTION OF NEW SPECIES.

group of the former one in another geological formation. It may, however, happen, that though later in time, the new series of species may never attain to so high a degree of organization as those preceding it, but in its turn become extinct, and give place to yet another modification from the same root, which may be of higher or lower organization, more or less numerous in species, and more or less varied in form and structure than either of those which preceded it. Again, each of these groups may not have become totally extinct, but may have left a few species, the modified prototypes of which have existed in each succeeding period, a faint memorial of their former grandeur and luxuriance. Thus every case of apparent retrogression may be in reality a progress, though an interrupted one: when some monarch of the forest loses a limb, it may be replaced by a feeble and sickly substitute. The foregoing remarks appear to apply to the case of the Mollusca, which, at a very early period, had reached a high organization and a great development of forms and species in the testaceous Cephalopoda. In each succeeding age modified species and genera replaced the former ones which had become extinct, and as we approach the present æra, but few and small representatives of the group remain, while the Gasteropods and Bivalves have acquired an immense preponderance. In the long series of changes the earth has undergone, the process of peopling it with organic beings has been continually going on, and whenever any of the higher groups have become nearly or quite extinct, the lower forms which have better resisted the modified physical conditions have served as the antitypes on which to found the new races. In this manner alone, it is believed, can the representative groups at successive periods, and the risings and fallings in the scale of organization, be in every case explained.

Objections to Forbes’ Theory of Polarity.

The hypothesis of polarity, recently put forward by Professor Edward Forbes to account for the abundance of generic forms at a very early

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