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nydus/Contributions to the Theory of Natural SelectionPublic

This volume collects essays written by Alfred Russel Wallace regarding his independent discovery and development of the theory of natural selection. The author provides corrected and expanded versions of his previous scientific writings to demonstrate the scope and application of his findings.

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Table of Contents

IV. THE MALAYAN PAPILIONIDÆ OR SWALLOW-TAILED BUTTERFLIES, AS ILLUSTRATIVE OF THE THEORY OF NATURAL SELECTION.

  1. Co-existing Variety.—This is a somewhat doubtful case. It is when a slight but permanent and hereditary modification of form exists in company with the parent or typical form, without presenting those intermediate gradations which would constitute it a case of simple variability. It is evidently only by direct evidence of the two forms breeding separately that this can be distinguished from dimorphism. The difficulty occurs in Papilio Jason, and P. Evemon, which inhabit the same localities, and are almost exactly alike in form, size, and colouration, except that the latter always wants a very conspicuous red spot on the under surface, which is found not only in P. Jason, but in all the allied species. It is only by breeding the two insects that it can be determined whether this is a case of a co-existing variety or of dimorphism. In the former case, however, the difference being constant and so very conspicuous and easily defined, I see not how we could escape considering it as a distinct species. A true case of co-existing forms would, I consider, be produced, if a slight variety had become fixed as a local form, and afterwards been brought into contact with the parent species, with little or no intermixture of the two; and such instances do very probably occur.
  1. Race or subspecies.—These are local forms completely fixed and isolated; and there is no possible test but individual opinion to determine which of them shall be considered as species and which varieties. If stability of form and “*the constant transmission of some characteristic
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