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nydus/Contributions to the Theory of Natural SelectionPublic

This volume collects essays written by Alfred Russel Wallace regarding his independent discovery and development of the theory of natural selection. The author provides corrected and expanded versions of his previous scientific writings to demonstrate the scope and application of his findings.

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Table of Contents

II. ON THE TENDENCY OF VARIETIES TO DEPART INDEFINITELY FROM THE ORIGINAL TYPE.

The Law of Population of Species.

The general proportion that must obtain between certain groups of animals is readily seen. Large animals cannot be so abundant as small ones; the carnivora must be less numerous than the herbivora; eagles and lions can never be so plentiful as pigeons and antelopes; and the wild asses of the Tartarian deserts cannot equal in numbers the horses of the more luxuriant prairies and pampas of America. The greater or less fecundity of an animal is often considered to be one of the chief causes of its abundance or scarcity; but a consideration of the facts will show us that it really has little or nothing to do with the matter. Even the least prolific of animals would increase rapidly if unchecked, whereas it is evident that the animal population of the globe must be stationary, or perhaps, through the influence of man, decreasing. Fluctuations there may be; but permanent increase, except in restricted localities, is almost impossible. For example, our own observation must convince us that birds do not go on increasing every year in a geometrical ratio, as they would do, were there not some powerful check to their natural increase. Very few birds produce less than two young ones each year, while many have six, eight, or ten; four will certainly be below the average; and if we suppose that each pair produce young only four times in their life, that will also be below the average, supposing them not to die either by violence or want of food. Yet at this rate how tremendous would be the increase in a few years from a single pair! A simple calculation will show that in fifteen years each pair of birds would have increased to nearly ten millions!1 whereas we have no reason to believe that the number of the birds of any country increases at all in fifteen or in one hundred and fifty years. With such powers of increase the population must have reached its limits, and have become stationary, in a very few years after the origin of each species. It is evident, therefore, that each year an immense number of birds must perish—as many in fact as are born; and as on the lowest calculation the progeny are each year twice as numerous as their parents,

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