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nydus/Contributions to the Theory of Natural SelectionPublic

This volume collects essays written by Alfred Russel Wallace regarding his independent discovery and development of the theory of natural selection. The author provides corrected and expanded versions of his previous scientific writings to demonstrate the scope and application of his findings.

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Table of Contents

II. ON THE TENDENCY OF VARIETIES TO DEPART INDEFINITELY FROM THE ORIGINAL TYPE.

unlike itself and resembling the other. This, however, would seem quite incompatible with the “permanent invariability of species,” but the difficulty is overcome by assuming that such varieties have strict limits, and can never again vary further from the original type, although they may return to it, which, from the analogy of the domesticated animals, is considered to be highly probable, if not certainly proved.

It will be observed that this argument rests entirely on the assumption, that varieties occurring in a state of nature are in all respects analogous to or even identical with those of domestic animals, and are governed by the same laws as regards their permanence or further variation. But it is the object of the present paper to show that this assumption is altogether false, that there is a general principle in nature which will cause many varieties to survive the parent species, and to give rise to successive variations departing further and further from the original type; and which also produces, in domesticated animals, the tendency of varieties to return to the parent form.

The Struggle for Existence.

The life of wild animals is a struggle for existence. The full exertion of all their faculties and all their energies is required to preserve their own existence and provide for that of their infant offspring. The possibility of procuring food during the least favourable seasons, and of escaping the attacks of their most dangerous enemies, are the primary conditions which determine the existence both of individuals and of entire species. These conditions will also determine the population of a species; and by a careful consideration of all the circumstances we may be enabled to comprehend, and in some degree to explain, what at first sight appears so inexplicable—the excessive abundance of some species, while others closely allied to them are very rare.

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