It follows from the experiments of M. Delage that it is this last indication which is much the most important; doubtless because the nerves are less sensible to the difference of pressure itself than to the brusque variations of this difference. The first three indications may thus be neglected.
Knowing the acceleration of the motion of rotation of the head at each instant, we deduce from it, by an unconscious integration, the final orientation of the head, referred to a certain initial orientation taken as origin. The circular canals contribute, therefore, to inform us of the movements that we have executed, and that on the same ground as the muscular sensations. When, therefore, above we speak of the series S or of the series Σ, we should say, not that these were series of muscular sensations alone, but that they were series at the same time of muscular sensations and of sensations due to the semicircular canals. Apart from this addition, we should have nothing to change in what precedes.
In the series S and Σ, these sensations of the semicircular canals evidently hold a very important place. Yet alone they would not suffice, because they can tell us only of the movements of the head; they tell us nothing of the relative movements of the body or of the members in regard to the head. And more, it seems that they tell us only of the rotations of the head and not of the translations it may undergo.