Fig. 58. Final stage in the first segôÙmenôÙtaôÙtion of the egg of CereôÙbratôÙuôÙlus. (From PreôÙnant, after Coe.) 245 Fig. 59. Diagram of field of force with two similar poles.
As we study these manifold phenomena, in the individual cases of particular plants and animals, we recognise a close identity of type, coupled with almost endless variation of specific detail; and in particular, the order of succession in which certain of the phenomena occur is variable and irregular. The precise order of the phenomena, the time of longitudinal and of transverse fission of the chromatin thread, of the break-up of the nuclear wall, and so forth, will depend upon various minor contingencies and ãinterferences.ã And it is worthy of particular note that these variations, in the order of events and in other subordinate details, while doubtless attributable to specific physical conditions, would seem to be without any obvious clasôÙsiôÙfiôÙcaôÙtory value or other biological significance48.
As regards the actual mechanical division of the cell into two halves, we shall see presently that, in certain cases, such as that of a long cylindrical filament, surface-tension, and what is known as the principle of ãminimal area,ã go a long way to explain the mechanical process of division; and in all cells whatsoever, the process of division must somehow be explained as the result of a conflict between surface-tension and its opposing forces. But in such a case as our spherical cell, it is not very easy to see what physical cause is at work to disturb its equiôÙlibôÙrium and its integrity.
The fact that, when actual division of the cell takes place, it does so at right angles to the polar axis and precisely in the direction of the equatorial plane, would lead us to suspect that the new surface formed in the equatorial plane sets up an annular tension, directed inwards, where it meets the outer surface layer of the cell itself. But at this point, the problem becomes more complicated. Before we could hope to comprehend it, we should have not only to enquire into the potential distribution at the surface of the cell in relation to that which we have seen to exist in its interior, but we should probably also have to take account of the differences