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CHAPTER VIII THE FORMS OF TISSUES OR CELL-AGGREGATES ( continued )

are not only stating empirically that the partition tends to appear in a small, rather than a large cross-section of the cell: but we are also implicitly ascribing to the cell a longitudinal polarity (Fig. 143, A), and implicitly asserting that it tends to {357} divide (just as the segmenting egg does), by a partition transverse to its polar axis. Such a polarity may conceivably be due to a chemical asymmetry, or anisotropy, such as we have learned of (from Professor Macallum〙s experiments) in our chapter on Adsorption. Now if the chemical concentration, on which this anisotropy or polarity (by hypothesis) depends, be unsymmetrical, one of its poles being as it were deflected to one side, where a little branch or bud is being (or about to be) given off,〔all in precise accordance with the adsorption phenomena described on p. 289,〔then our 〜polar axis〝 would necessarily be a curved axis, and the partition, being constrained (again ex hypothesi) to arise transversely to the polar axis, would lie obliquely to the apparent axis of the cell (Fig. 143, B, C). And if the oblique partition be so situated that it has to meet the opposite walls (as in C), then, in order to do so symmetrically (i.e. either perpendicularly, as when the cell-wall is already solidified, or at least at equal angles on either side), it is evident that the partition, in its course from one side of the cell to the other, must necessarily assume a more or less S-shaped curvature (Fig. 143, D).

As a matter of fact, while we have abundant simple illustrations of the principles which we have now begun to study, apparent exceptions to this simplicity, due to an asymmetry of the cell itself, or of the system of which the single cell is but a part, are by no means rare. For example, we know that in cambium-cells, division

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