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nydus/On Growth and FormPublic
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CHAPTER V THE FORMS OF CELLS

language, each particle is free to take up, and does take up, its position of least potential energy relative to those already deposited; in other words, for each particle motion is induced until the energy of the system is so distributed that no tendency or resultant force remains to move it more. The application of this principle has been shewn to lead to the production of planes3 (in all cases where by the limitation of material, surfaces must occur); and where we have planes, straight edges and solid angles must obviously also occur; and, if equiôÙlibôÙrium is {203} to follow, must occur symmetrically. Our piling up of shot, or manufacture of mimic crystals, gives us visible demonstration that the result is actually to obtain, as in the natural crystal, plane surfaces and sharp angles, symmetrically disposed.

But the living cell grows in a totally different way, very much as a piece of glue swells up in water, by 〜imbibition,〝 or by interpenetration into and throughout its entire substance. The semifluid colloid mass takes up water, partly to combine chemically with its individual molecules4, partly by physical diffusion into the interstices between these molecules, and partly, as it would seem, in other ways; so that the entire phenomenon is a very complex and even an obscure one. But, so far as we are concerned, the net result is a very simple one. For the equiôÙlibôÙrium or tendency to equiôÙlibôÙrium of fluid pressure in all parts of its interior while the process of imbibition is going on, the constant rearrangement of its fluid mass, the contrast in short with the crystalline method of growth where each particle comes to rest to move (relatively to the whole) no more, lead the mass of jelly to swell up, very much as a bladder into which we blow air, and so, by a graded and harmonious distribution of forces, to assume everywhere a rounded and more or less bubble-like external form5. So, when the same school of older naturalists called attention to a new distinction or contrast of form between the organic and inorganic objects, in that the contours of the former tended to roundness and curvature, and those of the latter to be bounded by straight lines, planes and sharp angles, we see that this contrast was not a new and different one, but only another aspect of their former statement, and an immediate consequence of the difference between the processes of agglutination and intussusception.

This common and general contrast between the form of the crystal on the one hand, and of the colloid or of the organism on the other, must by no means be pressed too far. For Lehmann, {204} in his great work on so-called Fluid Crystals6, to which we shall afterwards return, has shewn how, under certain circumstances, surface-tension phenomena may coexist with cryôÙstalôÙliôÙsaôÙtion, and produce a form of minimal potential which is a resultant of both: the fact being that the bonds maintaining the crystalline arrangement are now so much looser than in the solid condition that the tendency to least

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