Its substance will never be a perfect fluid, and hence equiôÙlibôÙrium will be more or less slowly reached; its surface will seldom be perfectly homogeneous, and therefore equiôÙlibôÙrium will (in the fluid condition) seldom be perfectly attained; it will very often, or generally, be the seat of other forces, symmetrical or unsymmetrical; and all these causes will more or less perturb the effects of surface tension acting by itself. But we shall find that, on the whole, these effects of surface tension though modified are not obliterated nor even masked; and accordingly the phenomena to which I have devoted the foregoing pages will be found manifestly recurring and repeating themselves among the phenomena of the organic cell.
In a spiderãs web we find exemplified several of the principles {232} of surface tension which we have now explained. The thread is formed out of the fluid secretion of a gland, and issues from the body as a semi-fluid cylinder, that is to say in the form of a surface of equiôÙlibôÙrium, the force of expulsion giving it its elongation and that of surface tension giving it its circular section. It is prevented, by almost immediate solidification on exposure to the air, from breaking up into separate drops or spherules, as it would otherwise tend to do as soon as the length of the cylinder had passed its limit of stability. But it is otherwise with the sticky secretion which, coming from another gland, is simultaneously poured over the issuing thread when it is to form the spiral portion of the web. This latter secretion is more fluid than the first, and retains its fluidity for a very much longer time, finally drying up after several hours. By capillarity it ãwetsã the thread, spreading itself over it in an even film, which film is now itself a cylinder. But this liquid cylinder has its limit of stability when its length equals its own circumference, and therefore just at the points so defined it tends to disrupt into separate segments: or rather, in the actual case, at points somewhat more distant, owing to the imperfect fluidity of the viscous film, and still more to the frictional drag upon it of the inner solid cylinder, or thread, with which it is in contact. The cylinder disrupts in the usual manner, passing first into the wavy outline of an unduloid, whose swollen portions swell more and more till the contracted parts break asunder, and we arrive at a series of spherical drops or beads, of equal size, strung at equal intervals along the thread. If we try to spread varnish over a thin stretched wire, we produce automatically the same identical result25; unless our varnish be such as to dry almost instantaneously, it gathers into beads, and do what we can, we fail to spread it smooth. It follows that, according to the viscidity and drying power of the varnish, the process may stop or seem to stop at any point short of the formation of the perfect spherules; it is quite possible, therefore, that as